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Publications

Scientific Articles:

Tim S Jessop, Deni Purwandana, M Jeri Imansyah, Claudio Ciofi, Y Jackson Benu, Achmad Ariefiandy (2022), The influence of tropical seasonality on breeding phenology, growth, survival and movement of a large reptile (Varanus komodoensis), Biological Journal of the Linnean Society, Volume 136, Issue 4, August 2022, Pages 552–565, https://doi.org/10.1093/biolinnean/blac045

Purwandana, D., Ariefiandy, A., Azmi, M., Nasu, S. A., Dos, A. A., & Jessop, T. S. (2021). Turning ghosts into dragons: improving camera monitoring outcomes for a cryptic low-density Komodo dragon population in eastern Indonesia. Wildlife Research.

Ariefiandy, A., Purwandana, D., Azmi, M., Panggur, M. R., Mardani, J., Parra, D.P., Jessop, T.S. (2021). Invasive water buffalo population trends and competition‑related consequences for native rusa deer in eastern Indonesian protected areas. Mammalian Biology. doi: 10.1007/s42991-021-00161-y

Azmi, M., Ardiantiono, Nasu, S. A., Kasim, A. M., Ariefiandy, A., Purwandana, D., Ciofi, C., Jessop, T. J. (2021). Incidences of road kills and injuries of komodo dragons along the North Coast of Flores Island, Indonesia. Herpetological Conservation and Biology, 16(1):11–16. 

Purwandana, D., Ciofi C., Imansyah, M. J., Ariefiandy A., Rudiharto H., Jessop, T.S. (2021). Prey preferences and body mass most influence movement behavior and home range area of Komodo Dragons. Ichthyology & Herpetology, 109(1), 92-101.

Purwandana, D., Imansyah, M. J., Ariefiandy, A., Rudiharto, H., Ciofi, C., & Jessop, T. S. (2020). Insights into the Nesting Ecology and Annual Hatchling Production of the Komodo Dragon. Copeia, 108(4), 855-862.

Jones, A. R., Jessop, T. S., Ariefiandy, A., Brook, B. W., Brown, S. C., Ciofi, C., … & Fordham, D. A. (2020) Identifying island safe havens to prevent the extinction of the World’s largest lizard from global warming. Ecology and Evolution.

Jessop,  T.  S.,  A.  Ariefiandy,  D.  M.  Forsyth,  D.  Purwandana,  C.  R.  White,  Y.  J.  Benu,  T.  Madsen,  H.  J.  Harlow,  and  M.  Letnic. (2020). Komodo dragons are not ecological analogs of apex mammalian predators. Ecology 101(4):e02970. 10.1002/ecy.2970

Kamil, P.I., Susianto, H., Purwandana, D., Ariefiandy, A. (2019). Anthropomorphic and factual approaches in Komodo dragon conservation awareness program for elementary school students: Initial study. Applied Environmental Education & Communication. doi: 10.1080/1533015X.2019.1582374.

Ariefiandy, A., Purwandana, D., Benu, Y. J., Letnic, M., Jessop, T.S. (2019). Knee deep in trouble: rusa deer use an aquatic escape behaviour to delay attack by Komodo dragons. Australian Mammalogy. doi: 10.1071/AM18052.

Jessop, T. S., Ariefiandy, A., Purwandana, D., Benu, Y. J., Hyatt, M., & Letnic, M. (2019). Little to fear: largest lizard predator induces weak defense responses in ungulate prey. Behavioral Ecology.

Jessop, T., Ariefiandy, A., Purwandana, D., Ciofi, C., Imansyah, J., Benu, Y. J., Fordham, D. A., Forsyth, D. M., Mulder, R. A., Phillips, B. L. (2018). Exploring mechanisms and origins of reduced dispersal in islands Komodo dragons. Proc. R. Soc. B 285: 20181829.

Ariefiandy, A., Purwandana, D., Nasu, S.A., Imansyah, M.J., Benu, Y. J., Chrismiawati, M., Kamil, P.I.,  Ciofi, C. and Jessop, T. (2017). Panduan Lapangan Biawak Komodo (Komodo Dragon Field Guide), Yayasan Komodo Survival Program, Denpasar.

Imansyah, M.J., Purwandana, D., Ariefiandy, A., Benu, Y. J., Jessop, T., & Trainor, C. R. (2016). Valley-floor censuses of the Critically Endangered Yellow-crested Cockatoo Cacatua sulphurea occidentalis on Komodo Island, East Nusa Tenggara province, Indonesia, point to a steep population decline over a six-year period. Forktail, 32: 66 – 71.

Ariefiandy, A., Forsyth, D.M., Purwandana, D., Imansyah, J., Ciofi, C., Rudiharto, H., Seno, A. and Jessop, T.S. (2016) Temporal and spatial dynamics of insular Rusa deer and wild pig populations in Komodo National Park. Journal of Mammalogy: p.gyw131.

Purwandana, D., Ariefiandy, A., Imansyah, M.J., Seno, A., Ciofi, C., Letnic, M. and Jessop, T.S., (2016). Ecological allometries and niche use dynamics across Komodo dragon ontogeny. The Science of Nature, 103(3-4):1-11.

Ariefiandy, A., Purwandana D., Natali, C., Imansyah, M. J., Surahman, M., Jessop, T. S. & Ciofi, C. (2015) Conservation of Komodo dragons Varanus komodoensis in the Wae Wuul nature reserve, Flores, Indonesia: a multidisciplinary approach. International Zoo Yearbook, 49: 67–80. doi: 10.1111/izy.12072.

Ariefiandy, A., Purwandana D., Nasu, S. A., Surahman, M., Ciofi, C. & Jessop, T. S. (2015) First record of Komodo dragon nesting activity and hatchling emergence from north Flores, eastern Indonesia. Biawak, 9:33-35.

Purwandana, D., Ariefiandy, A., Imansyah, M. J., Ciofi, C., Forsyth, D. M., Gormley, A. M., Rudiharto, H., Seno, A., Fordham, D. A., Gillespie, G. and Jessop, T. S. (2015), Evaluating environmental, demographic and genetic effects on population-level survival in an island endemic. Ecography, 38: 1060–1070. doi: 10.1111/ecog.01300.

Ariefiandy, A., Purwandana D., Seno, A., Chrismiawati, M., Jessop T.S. & Ciofi, C. (2014) Evaluation of three field monitoring-density estimation protocols and their relevance to Komodo dragon conservation. Biodiversity and Conservation. Published online, 01 July 2014. doi:10.1007/s10531-014-0733-3.

Purwandana D., Ariefiandy, A., Imansyah, M. J., Rudiharto, H., Seno, A., Ciofi C., Fordham, D. A. & Jessop T.S. (2014) Demographic status of Komodo dragon populations in Komodo National Park. Biological Conservation, 171, 29-35.

Ariefiandy, A., Purwandana D., Aganto S., Ciofi C. and Jessop T.S. (2013) Can camera traps monitor Komodo dragons a large ectothermic predator? PLoS ONE 8(3): e58800. doi:10.1371/journal.pone.0058800.

Ariefiandy, A., Purwandana, D.,  Coulson, G., Forsyth, D. M. and Jessop, T. S. (2013). Monitoring the ungulate prey of the Komodo dragon: distance sampling or faecal counts? Wildlife Biology 19:1-12.

Jessop, T., Sumner, J., Imansyah, J., Purwandana, D., Seno, A., Ariefiandy, A., Ciofi, C. (2010). Assessment of environmental and host dependent factors correlated with tick abundance on komodo dragons. Australian Zoologist 35(2), pp. 265-275.

Jessop, T., Imansyah, J., Purwandana, D., Ariefiandy, A., Opat, D. S. (2008). Incidence of fish hook ingestion by komodo dragons. Biawak 2(3): 121-123.

Ciofi, C., de Boer, M. (2004). Distribution and conservation of the komodo monitor (Varanus komodoensis). Herpetological Journal 14(2): 99-107.

Ciofi, C., Puswati, J., Winana, D., De Boer, M. E., Chelazzi, G., and Sastrawan, P. (2007). Preliminary analysis of home range structure in the Komodo monitor, Varanus komodoensis. Copeia 2: 462-470.

Ciofi, C., Beaumont, M.A., Swingland, I.R., Bruford, M.W. (1999). Genetic divergence and units for conservation in the Komodo dragon Varanus komodoensis. Proc. R. Lond. B, 266: 2269-2274.

Ciofi, C., Bruford, M.W. (1999). Genetic structure and gene flow among Komodo dragon populations inferred by microsatellite loci analysis. Molecular Ecology 8: S17-S30.

Imansyah, M. J., Jessop, T. S., Sumner, J., Purwandana, D.,  Ariefiandy, A. and Seno,. A. (2009) Distribution, seasonal use, and predation of incubation mounds of Orange-footed Scrubfowl on Komodo Island, Indonesia. J. Field Ornithol. 80: 119–126.

Imansyah, M. J., Jessop, T. S., Ciofi, C., and Akbar, Z. (2008). Ontogenetic differences in the spatial ecology of immature Komodo dragons. Journal of Zoology 274, 107-115.

Jessop, T.S., Madsen, T., Ciofi, C., Imansyah, M.J., Purwandana, D., Rudiharto, H., Arifiandy, A., Phillips, J.A. (2007). Island differences in population size structure and catch per unit effort and their conservation implications for Komodo dragons. Biological Conservation 135:247-255.

Jessop, T.S., Madsen, T., Sumner J., Rudiharto, H., Phillips, J.A., Ciofi, C. (2006). Maximum body size among insular Komodo dragon populations covaries with large prey density. Oikos 112: 422-429.

Jessop T.S., Sumner J., Rudiharto H., Purwandana D., Imansyah M.J. & Phillips, J.A. (2004). Distribution, use and selection of nest type by Komodo Dragons. Biological Conservation 117: 463 – 470.

Harlow, H.J., Purwandana, D., Jessop, T.S., Phillips, J.A. (2010). Body temperature and thermoregulation of komodo dragons in the field. Journal of Thermal Biology 35(7): 338-347.

Harlow, H.J., Purwandana, D., Jessop, T.S., Phillips, J.A. (2010). Size-related differences in the thermoregulatory habits of free-ranging komodo dragons. International Journal of Zoology, DOI: 10.1155/2010/921371.

Laver, R.J., Purwandana, D., Ariefiandy, A., Imansyah, J., Forsyth, D., Ciofi, C., Jessop, T.S. (2012). Life-history and spatial determinants of somatic growth dynamics in komodo dragon populations. PLoS ONE 7(9): e45398.

Thesis:

Ariefiandy, A. (2011). Population assessments of ungulate prey and Komodo dragons across protected areas in eastern Indonesia. M.Phil. thesis. The University of Melbourne, Melbourne.

Imansyah, M.J. (2006). Spatial ecology of hatchling and juvenile Komodo dragons in the Komodo National Park, Indonesia. MSc thesis. University Kebangsaan Malaysia, Bangi.

Purwandana, D., (2007). Nesting activity and spatial ecology of female komodo dragons (Varanus komodoensis) in the komodo national park, Indonesia. MSc thesis. University Kebangsaan Malaysia, Bangi.

Report:

Ariefiandy, A., Purwandana, D.,  Seno, A.,  Ciofi, C. dan Jessop, T. S. (2012).  Penggunaan camera trap untuk pemantauan populasi biawak Komodo di Taman Nasional Komodo, Indonesia. Laporan dari Komodo Survival Program dan Balai Taman Nasional Komodo, Labuan Bajo, Flores.

Imansyah, M.J., Anggoro, D.G., Yangpatra, N., Hidayat, A. dan Benu, Y.J. (2005). Sebaran dan karakteristik pohon sarang kakatua jambul kuning (Cacatua sulphurea parvula) di Pulau Komodo,

Taman Nasional Komodo. Laporan dari the Zoological Society of San Diego, Balai Taman Nasional Komodo, dan The Nature Conservancy. Labuan Bajo, Flores. Labuan Bajo, Flores.

Imansyah, M.J., Purwandana, D., Rudiharto, H.,dan Jessop, T.S. (2003). Survey potensi hidupan liar terestrial di Pulau KOmodo, taman Nasional Komodo tahun 2002. Laporan dari the Zoological Society of San Diego, Balai Taman Nasional Komodo, dan The Nature Conservancy. Labuan Bajo, Flores.

Imansyah, M.J., Purwandana, D. dan Jessop, T.S. (2002). Materi Kursus 1 Dasar-dasar Sistem Informasi Geografis untuk Staff Taman Nasional Komodo. Laporan dari the Zoological Society of San Diego, Balai Taman Nasional Komodo, dan The Nature Conservancy. Labuan Bajo, Flores.

Jessop, T.S, Imansyah, M.J, Purwandana, D., Ariefiandy, A., Rudiharto, H., Seno, A., Opat D.S., Noviandi, T., Payung, I., Ciofi, C. (2007). Ekologi populasi, reproduksi, dan spasial biawak Komodo (Varanus komodoensis) di Taman Nasional Komodo. Disunting oleh Imansyah, M.J., Ariefiandy, A.,

Purwandana, D. Laporan dari the Zoological Society of San Diego, Balai Taman Nasional Komodo, dan The Nature Conservancy. Labuan Bajo, Flores.

Jessop, T.S., Imansyah, M.J., Purwandana, D., , Arifiandy, A., Rudiharto, H.  (2007). Panduan Pemantauan Ekologi dan Hidupan Liar di TN Komodo, Indonesia. Laporan dari the Zoological Society of San Diego, Balai Taman Nasional Komodo, dan The Nature Conservancy. Labuan Bajo, Flores.

Jessop, T.M., Sumner, J., Imansyah, M.J., Purwandana, D., Ariefiandy, A., & Seno, A., (2006). Penilaian Distribusi, Penggunaan Musiman, dan Predasi Sarang Burung Gosong – Kaki Merah di Pulau Komodo. Laporan terjemahan berbahasa Indonesia. Laporan dari the Zoological Society of San Diego, Balai Taman Nasional Komodo, dan The Nature Conservancy. Labuan Bajo, Flores.

Jessop, T.S., Forsyth, D.M., Purwandana, D., Seno, A. (2006). Efektivitas Penggunaan Metode Penghitungan Kotoran Untuk Menilai Distribusi dan Kelimpahan Relatif Monyet dan Musang di Taman Nasional Komodo, Indonesia. Terjemahan. Ariefiandy, A., Purwandana, D. Imansyah, M.J. Laporan dari the Zoological Society of San Diego, Balai Taman Nasional Komodo, dan The Nature Conservancy. Labuan Bajo, Flores, Indonesia.

Jessop, T.S., Forsyth, D.M., Purwandana, D., Imansyah, M.J., Opat, D.S., and McDonald-Madden, E. (2005). Monitoring the ungulate prey of komodo dragons (Varanus komodoensis) using faecal counts. Report from the Zoological Society of San Diego, Komodo National Park, and The Nature Conservancy. Labuan Bajo, Flores.

Jessop, T.S., Forsyth, D.M., Purwandana, D., Imansyah, M.J., Opat, D.S., dan McDonald-Madden, (2005). Pemantauan mangsa ungulata biawak komodo (Varanus komodoensis) dengan menggunakan metode penghitungan kotoran. Terjemahan. Ariefiandy, A., Purwandana, D. Imansyah, M.J. Laporan dari the Zoological Society of San Diego, Balai Taman Nasional Komodo, dan The Nature Conservancy. Labuan Bajo, Flores.

Jessop, T.S., Madsen, T., Sumner., J., Rudiharto, H., Phillips, J.A. dan Ciofi, C. (2005). Ukuran tubuh-maksimum antar populasi-terbatas-pulau biawak Komodo dan keterkaitannya dengan kepadatan mangsa besar. Laporan dari the Zoological Society of San Diego, Balai Taman Nasional Komodo, dan The Nature Conservancy. Labuan Bajo, Flores.

Jessop, T.S., Madsen, T., Purwandana, D., Imansyah, M.J., Rudiharto, H. and Ciofi, C. (2005). Evidence for energetic constraints affecting a small island Komodo dragon population. Report from the Zoological Society of San Diego, Komodo National Park, and The Nature Conservancy. Labuan Bajo, Flores.

Jessop T.S., Sumner J., Rudiharto H., Phillips, J.A., and Ciofi, C. (2004). Variasi ukuran tubuh antar pupulasi terbatas-pulau pada Komodo (Varanidae). Laporan dari the Zoological Society of San Diego, Balai Taman Nasional Komodo, dan The Nature Conservancy. Labuan Bajo, Flores.

Jessop, T.S., Sumner J., Rudiharto H., Purwandana D., Imansyah M.J. & Phillips, J.A. (2003). Studi Distribusi, Penggunaan dan Pemilihan Tipe Sarang oleh Biawak Komodo: Implikasi untuk Konservasi dan Manajemen, Laporan dari the Zoological Society of San Diego, Balai Taman Nasional Komodo, dan The Nature Conservancy. Labuan Bajo, Flores.

Jessop, T.S., Sumner J., Rudiharto H., Purwandana D., Imansyah M.J., (2002). Kursus Metode Sampling dan Statistik dalam Populasi Tertutup Digunakan untuk Penaksiran Kelimpahan Varanus Komodoensis. Laporan dari the Zoological Society of San Diego, Balai Taman Nasional Komodo, dan The Nature Conservancy. Labuan Bajo, Flores.

Further Readings:

Barnard, T.P. (2011). Protecting the dragon: Dutch attempts at limiting access to komodo lizards in the 1920s and 1930s. Indonesia, 92: 97-123.

D’Amore, D.C. (2011). The effects of biting and pulling on the forces generated during feeding in the komodo dragon (Varanus komodoensis). PLoS ONE 6(10): e26226.

Bull, J.J., Jessop, T.S., Whiteley, M. (2010). Deathly drool: Evolutionary and ecological basis of septic bacteria in komodo dragon mouths. PLoS ONE 5(6): e11097.

Fry, B.G., Wroe, S., Teeuwisse, W., van Osch, M.J.P., Moreno, K., …, Norman, J.A. (2009). A central role for venom in predation by Varanus komodoensis (komodo dragon) and the extinct giant Varanus (Megalania) priscus. PNAS 106(22): 8969-8974.

Sunter, G. (2008). Management and reproduction of the Komodo dragon Varanus komodoensis Ouwens 1912 at ZSL London Zoo. International Zoo Yearbook, DOI: 10.1111/j.1748-1090.2007.00029.x

Montgomery, J.M., Gillespie, D., Sastrawan, P., Fredeking, T.M., Stewart, G.L. (2002). Aerobic salivary bacteria in wild and captive komodo dragons. J Wild Dis. 38(3): 545-551.

Walpole, M.J. (2001). Feeding dragons in Komodo National Park: a tourism tool with conservation complications. Animal Conservation 4(1): 67-73.

Gillespie, D., Frye, F.L., Stockham, S.L., Fredeking, T. (2000). Blood values in wild and captive Komodo dragons (Varanus komodoensis). Zoo Biol 19(6): 495-509.

Green, B., King, D., Braysher, M., Saim, A. (1991). Thermoregulation, water turnover and energetics of free-living komodo dragons, Varanus komodoensis. Comparative Biochemistry and Physiology Part A: Physiology 99(1-2): 97-101.

Diamond, J.M. (1987). Did Komodo dragons evolve to eat pygmy elephants?. Nature: No. 326.

Burden, W.D. (1928). Results of the Douglas Burden Expedition to the Island of Komodo. American Museum Novitates: No. 316.

Dunn, E. R. (1927). ‘Results of the Douglas Burden Expedition to the Island of Komodo. I. Notes on Varanus komodoensis.’ American Museum Novitates: No. 286.